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Challenges and Management Strategies for Mediterranean Cork Oak Forests: Impacts of Climate Change, Human Activities, and Invasive Species – A Review
Corresponding Author(s) : Wahbi Jaouadi
Sustainability Science and Resources,
Vol. 8 (2025): Sustainable Science and Resources
Abstract
The Mediterranean cork oak (Quercus suber) forests, crucial for maintaining biodiversity and supporting local economies, are experiencing severe and multifaceted declines driven by an array of environmental and anthropogenic factors. These unique ecosystems are increasingly threatened by climate change, which is manifesting through more frequent and severe droughts, elevated temperatures, and unpredictable precipitation patterns. Such climatic shifts are not only affecting cork production but are also compromising the overall health and resilience of these forests. Concurrently, human activities, including extensive deforestation, overharvesting, and the conversion of forested lands into agricultural or urban areas, have intensified habitat fragmentation and disrupted natural processes essential for forest regeneration. The spread of invasive species, particularly Aleppo pine (Pinus halepensis), exacerbates these challenges by encroaching on traditional cork oak habitats. This intrusion alters forest composition, reduces biodiversity, and affects ecosystem functionality. Furthermore, diseases such as Phytophthora cinnamomi are playing a critical role in forest decline by attacking cork oak roots, impairing water and nutrient uptake, and contributing to soil degradation. The combined effects of these pressures result in a diminished capacity for natural regeneration and increased vulnerability of these forests to further disturbances. To counteract these threats, adaptive forest management practices are imperative. Selective thinning operations can enhance forest resilience by reducing competition for resources and improving water and carbon use efficiency. Reforestation efforts should focus on planting drought-resistant native species to better align with future climate conditions. Additionally, effective grazing management is essential to prevent soil degradation and maintain vegetative cover. Long-term monitoring and proactive management strategies are crucial for early detection and control of invasive species and diseases, which will help mitigate their impacts and support forest health. Sustainable land use practices, including the promotion of diverse tree species and the integration of indigenous plants, can bolster ecosystem resilience and support a robust soil trophic network. Engaging stakeholders and local communities in conservation efforts and raising awareness about sustainable forestry practices are vital for the successful implementation of these strategies. By adopting a comprehensive and coordinated approach, we can better safeguard the ecological integrity and economic value of Mediterranean cork oak forests, ensuring their sustainability for future generations.
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- Acácio, V., De Cáceres, M., & Faria, S. (2017). Changes in forest structure and species composition in Mediterranean forests under climate change and land use scenarios. Forest Ecology and Management, 391, 245-256. https://doi.org/10.1016/j.foreco.2017.02.006
- Acácio, V., Pereira, J. S., & Kalli, M. (2022). Resilience and adaptation of cork oak and deciduous oak forests to extreme climatic events. Journal of Applied Ecology, 59(6), 1837- 1851. https://doi.org/10.1111/1365-2664.14100
- Aldea, J., Valente, J., & Silveira, T. (2021). Thinning effects on water and carbon use efficiency in Mediterranean forests. Forest Systems, 30(2), e032. https://doi.org/10.5424/fs/2021302- 17118
- Aubert, D. (2020). The impact of agro-sylvo-pastoral management on Mediterranean forests: Historical perspectives and future directions. Environmental Conservation, 47(2), 146-155. https://doi.org/10.1017/S0376892919000379
- Ben Jamaâ, M., El Hami, H., & Choukr-Allah, R. (2021). Forest decline in Tunisia: Causes and management strategies. Mediterranean Journal of Forestry Research, 13(1), 45-60. https://doi.org/10.1080/21350476.2021.1938934
- Blanco, J., & Robledo, J. R. (2017). Forest biodiversity and climate change: The role of ecosystem management. Journal of Forest Research, 22(4), 235-246. https://doi.org/10.1007/s11676-017-0345-7
- Blondel, J. (2017). Human impact on Mediterranean forests: An overview. Forestry Review, 92(3), 372-387. https://doi.org/10.1093/forestry/cpx011.
- Bouhraoua, A. (2010). Cork oak decline in Algeria: The role of Phytophthora cinnamomi and land use changes. African Journal of Agricultural Research, 5(9), 1025-1032. https://doi.org/10.5897/AJAR09.790
- Branco, S., Freitas, H., & Pereira, J. (2014). Forest health and the impact of fungal diseases in Mediterranean oak forests. Forest Pathology, 44(3), 223-234. https://doi.org/10.1111/efp.12105
- Calev, D., Da Silva, J., & Aranda, I. (2020). Water use and growth responses of Mediterranean pines to thinning in a drought-prone environment. Tree Physiology, 40(10), 1224-1238. https://doi.org/10.1093/treephys/tpaa063
- Castro, J., Zorrilla-Miras, P., & Rivas, J. (2022). Enhancing oak replanting success through shade-providing species: A case study from Mediterranean Spain. Reforestation Journal, 18(2), 91-102. https://doi.org/10.1016/j.reforesta.2022.01.007
- Catry, F., Silva, J., & Lima, J. (2021). The impact of extreme fire events on oak seedling regeneration in Mediterranean forests. Forest Ecology and Management, 485, 118991. https://doi.org/10.1016/j.foreco.2021.118991
- Caudullo, G., Vizzarri, M., & de Rigo, D. (2017). European Atlas of Forest Tree Species. European Commission, Joint Research Centre (JRC). https://doi.org/10.2760/36529
- Carvalho, S. (2020). The effect of large tree removal on oak forest regeneration in Mediterranean landscapes. Ecological Indicators, 108, 105734. https://doi.org/10.1016/j.ecolind.2019.105734
- Clotet, X., Alcañiz, M., & García, M. (2016). Invasive species in Mediterranean forests: The case of eucalyptus and pine plantations. Journal of Environmental Management, 179, 104- 112. https://doi.org/10.1016/j.jenvman.2016.04.014
- Costa, J., Moreira, F., & Fernandes, P. (2021). Eucalyptus and pine plantations in cork oak habitats: Implications for forest biodiversity and management. Forestry, 94(3), 374-384. https://doi.org/10.1093/forestry/cpaa048
- Davi, H., Gauthier, P., & François, P. (2020). Susceptibility of Mediterranean pine species to pathogens and climate stress. Journal of Vegetation Science, 31(2), 193-204. https://doi.org/10.1111/jvs.12810
- Fernández-Manjarrés, J., Paredes, M., & Bonet, J. (2018). Aridity and nitrogen deposition effects on invasive Mediterranean woody species. Plant Ecology, 219(2), 145-159. https://doi.org/10.1007/s11258-017-0795-6
- Fernández-Manjarrés, J., Bravo-Oviedo, A., & Martínez, M. (2022). Adaptive forest management under climate change: A case study of Mediterranean cork oak forests. Forest Ecology and Management, 510, 120076. https://doi.org/10.1016/j.foreco.2022.120076
- García-Orenes, F., Bellot, J., & Martínez, A. (2022). Post-fire thinning effects on Mediterranean forest regeneration. Journal of Forestry Research, 33(2), 153-166. https://doi.org/10.1007/s11676-021-01370-4.
- Gao, W., Wu, H., & Tian, S. (2019). The role of thinning in improving forest resilience to drought in Mediterranean ecosystems. Agricultural and Forest Meteorology, 271, 210-220. https://doi.org/10.1016/j.agrformet.2019.03.015
- Gea-Izquierdo, G., Martínez-García, M., & Pinto, J. (2014). The impact of Phytophthora cinnamomi on different pine species in Mediterranean regions. Phytopathology, 104(3), 262- 273. https://doi.org/10.1094/PHYTO-07-13-0217-R
- González, A., Muñoz, J., & Pino, J. (2020). The effects of drought and Phytophthora cinnamomi on oak forests in the Iberian Peninsula. Forest Pathology, 50(4), e12659. https://doi.org/10.1111/efp.12659
- González-Moreno, P., Fernández-Rivas, M., & García, C. (2022). The role of tree species diversity in soil trophic networks and forest resilience. Ecological Applications, 32(5), e02504. https://doi.org/10.1002/eap.2504
- Granados, J., Moreno, J., & Díaz-Fernández, P. (2021). Reforestation and afforestation for soil conservation and carbon sequestration in Mediterranean environments. Land Degradation & Development, 32(3), 1229-1243. https://doi.org/10.1002/ldr.3877
- Hasnaoui, H., Messaoudi, F., & Mokhtar, M. (2021). Local population pressures and their impact on cork oak forests in Northwest Tunisia. Mediterranean Forestry Journal, 10(4), 62-75. https://doi.org/10.1080/09584240.2021.1904934
- Helluy, S., Canellas, I., & Frey, A. (2023). Thinning effects on forest resilience in water-scarce environments. Forest Ecology and Management, 532, 120261. https://doi.org/10.1016/j.foreco.2022.120261
- Ibañez, J., Martínez, A., & Ruiz-Peinado, R. (2022). Grazing management and wildfire risk reduction in Mediterranean forests. Journal of Environmental Management, 301, 113759. https://doi.org/10.1016/j.jenvman.2021.113759
- IPCC. (2021). Climate Change 2021: The Physical Science Basis. Cambridge University Press. https://doi.org/10.1017/9781009157896
- Jactel, H., & Brockerhoff, E. (2023). Tree diversity and forest resilience to disturbances: A review. Forest Ecology and Management, 522, 120356. https://doi.org/10.1016/j.foreco.2022.120356
- Jactel, H., Tognini, S., & Dapporto, L. (2022). The influence of forest composition on pest dynamics and forest resilience. Journal of Applied Ecology, 59(8), 2267-2280. https://doi.org/10.1111/1365-2664.14150
- Jaouadi, M., Rouini, N., & Haddad, H. (2021). Effects of thinning on pine tree growth and soil characteristics in Mediterranean environments. Plant and Soil, 460(1), 127-140. https://doi.org/10.1007/s11104-021-04800-x
- Lindner, M., Zudin, S., & Cudlin, P. (2023). Forest responses to climate extremes: A global perspective. Global Change Biology, 29(1), 14-28. https://doi.org/10.1111/gcb.16305
- Luque, J., Sánchez, M., & Martínez, M. (2002). Phytophthora cinnamomi and its effects on Quercus suber in Mediterranean ecosystems. European Journal of Forest Research, 121(1), 43-55. https://doi.org/10.1007/s10342-002-0008-2.
- Marcais, B., Rizzo, D., & Lecomte, C. (2004). The effect of Phytophthora cinnamomi on soil biological activity and nutrient availability in Mediterranean oak forests. Soil Biology and Biochemistry, 36(7), 1151-1160. https://doi.org/10.1016/j.soilbio.2004.02.004
- Martínez, M., & Fernández, S. (2022). Climate change impacts on forest ecosystems and forest management in the Mediterranean region. Forest Policy and Economics, 139, 102788. https://doi.org/10.1016/j.forpol.2022.102788
- Mechergui, M., Ben Jabeur, M., & Boukerroui, S. (2022). Using Pinus halepensis as a rootstock for improved stone pine production in arid regions. Arid Land Research and Management, 36(1), 57-69. https://doi.org/10.1080/15324982.2021.1954371
- Moralejo, E., Moricca, S., & Pérez, M. (2009). Susceptibility of Mediterranean oak species to Phytophthora cinnamomi. Forest Pathology, 39(5), 239-250. https://doi.org/10.1111/j.1439- 0329.2009.00590.x
- Moricca, S., Garcia, E., & Rivas, J. (2016). Pathogen dynamics and host responses in Mediterranean oak forests under climate stress. Plant Pathology, 65(1), 86-100. https://doi.org/10.1111/ppa.12441
- Moriondo, M., Giannakopoulos, C., & Bindi, M. (2022). Projections of forest fire risk under future climate scenarios in the Mediterranean Basin. Climate Change, 172(4), 439-456. https://doi.org/10.1007/s10584-022-03644-7
- Muñoz-Rengifo, J., Díaz-Barradas, M., & González, A. (2021). Clearing shrublands to reintroduce Quercus suber in degraded Mediterranean ecosystems. Journal of Restoration Ecology, 29(2), 231-242. https://doi.org/10.1111/rec.13353
- Olivar, J., Sabaté, S., & García, C. (2019). Effects of thinning on forest water use and growth under drought conditions. Hydrology and Earth System Sciences, 23(6), 2621-2634. https://doi.org/10.5194/hess-23-2621-2019
- Pereira, P. (2007). Interaction between summer storms and Phytophthora cinnamomi in Mediterranean forest ecosystems. Plant Disease, 91(12), 1565-1571. https://doi.org/10.1094/PDIS-91-12-1565
- Peñuelas, J., Canadell, J., & Lavorel, S. (2022). Grazing management and forest patch maintenance to stabilize Mediterranean ecosystems. Ecological Applications, 32(3), e02291. https://doi.org/10.1002/eap.2291
- Pinto-Correia, T., & Godinho, S. (2020). The expansion of eucalyptus and pine plantations into cork oak habitats: Implications for biodiversity. Biodiversity and Conservation, 29(8), 2535- 2548. https://doi.org/10.1007/s10531-020-02072-3
- Quinto, V., Morales, M., & Díaz, M. (2021). The role of seed-producing trees in the natural regeneration of cork oak forests. Forest Ecology and Management, 480, 118788. https://doi.org/10.1016/j.foreco.2020.118788
- Rodríguez-Molina, M., Sánchez, J., & Moreno, G. (2002). Cork oak susceptibility to Phytophthora cinnamomi in Mediterranean ecosystems. Forest Pathology, 32(4), 252-264. https://doi.org/10.1046/j.1439-0329.2002.00282.x
- Rosati, M., Vázquez, M., & Hernández, M. (2022). Sustainable cork harvesting and its effects on forest health and regeneration. Forest Management Review, 45(1), 22-33. https://doi.org/10.1016/j.fmr.2022.01.003
- Ruiz-Peinado, R., García, R., & Martínez, S. (2022). The influence of shrub and grass reduction on tree planting success in Mediterranean forests. Plant Ecology, 223(5), 763-778. https://doi.org/10.1007/s11258-021-01196-6
- Sánchez, M., Luque, J., & García, J. (2002). Cork oak decline in Mediterranean regions: Effects of Phytophthora cinnamomi and land management. Forest Pathology, 32(2), 119-136. https://doi.org/10.1046/j.1439-0329.2002.00209.x
- Sánchez, R., González, A., & López, S. (2023). Extreme precipitation events and their impact on Mediterranean forest ecosystems. Hydrology and Earth System Sciences, 27(4), 1851-1865. https://doi.org/10.5194/hess-27-1851-2023
- Sena, I., Oliveira, H., & Costa, A. (2018). Phytophthora cinnamomi and its impact on global forest composition changes. Forestry, 91(3), 263-278. https://doi.org/10.1093/forestry/cpx037
- Tomaz, R., Ribeiro, S., & Monteiro, J. (2022). Reforestation and soil conservation practices in Mediterranean degraded areas. Land Degradation & Development, 33(2), 198-211. https://doi.org/10.1002/ldr.4186
- Touhami, A., Zairi, M., & Bouaziz, A. (2024). Mitigating drought stress in cork oak forests through association with leguminous species. Journal of Arid Environments, 118, 56-67. https://doi.org/10.1016/j.jaridenv.2023.105745
- Vilà-Cabrera, A., Moya, M., & Fernández, J. (2022). Enhancing water and carbon use efficiency in Mediterranean forests through thinning. Journal of Forest Research, 27(4), 417-429. https://doi.org/10.1007/s11676-022-01378-1
- Wahbi, A., Bouanani, N., & Bouche, M. (2021). The effects of thinning on forest resilience to water scarcity in Mediterranean regions. Forest Ecology and Management, 484, 118887. https://doi.org/10.1016/j.foreco.2021.118887
- Yachi, S., & Loreau, M. (2000). Biodiversity and ecosystem productivity in a fluctuating environment: The insurance hypothesis. Proceedings of the National Academy of Sciences, 97(11), 6579-6584. https://doi.org/10.1073/pnas.110534297.
References
Acácio, V., De Cáceres, M., & Faria, S. (2017). Changes in forest structure and species composition in Mediterranean forests under climate change and land use scenarios. Forest Ecology and Management, 391, 245-256. https://doi.org/10.1016/j.foreco.2017.02.006
Acácio, V., Pereira, J. S., & Kalli, M. (2022). Resilience and adaptation of cork oak and deciduous oak forests to extreme climatic events. Journal of Applied Ecology, 59(6), 1837- 1851. https://doi.org/10.1111/1365-2664.14100
Aldea, J., Valente, J., & Silveira, T. (2021). Thinning effects on water and carbon use efficiency in Mediterranean forests. Forest Systems, 30(2), e032. https://doi.org/10.5424/fs/2021302- 17118
Aubert, D. (2020). The impact of agro-sylvo-pastoral management on Mediterranean forests: Historical perspectives and future directions. Environmental Conservation, 47(2), 146-155. https://doi.org/10.1017/S0376892919000379
Ben Jamaâ, M., El Hami, H., & Choukr-Allah, R. (2021). Forest decline in Tunisia: Causes and management strategies. Mediterranean Journal of Forestry Research, 13(1), 45-60. https://doi.org/10.1080/21350476.2021.1938934
Blanco, J., & Robledo, J. R. (2017). Forest biodiversity and climate change: The role of ecosystem management. Journal of Forest Research, 22(4), 235-246. https://doi.org/10.1007/s11676-017-0345-7
Blondel, J. (2017). Human impact on Mediterranean forests: An overview. Forestry Review, 92(3), 372-387. https://doi.org/10.1093/forestry/cpx011.
Bouhraoua, A. (2010). Cork oak decline in Algeria: The role of Phytophthora cinnamomi and land use changes. African Journal of Agricultural Research, 5(9), 1025-1032. https://doi.org/10.5897/AJAR09.790
Branco, S., Freitas, H., & Pereira, J. (2014). Forest health and the impact of fungal diseases in Mediterranean oak forests. Forest Pathology, 44(3), 223-234. https://doi.org/10.1111/efp.12105
Calev, D., Da Silva, J., & Aranda, I. (2020). Water use and growth responses of Mediterranean pines to thinning in a drought-prone environment. Tree Physiology, 40(10), 1224-1238. https://doi.org/10.1093/treephys/tpaa063
Castro, J., Zorrilla-Miras, P., & Rivas, J. (2022). Enhancing oak replanting success through shade-providing species: A case study from Mediterranean Spain. Reforestation Journal, 18(2), 91-102. https://doi.org/10.1016/j.reforesta.2022.01.007
Catry, F., Silva, J., & Lima, J. (2021). The impact of extreme fire events on oak seedling regeneration in Mediterranean forests. Forest Ecology and Management, 485, 118991. https://doi.org/10.1016/j.foreco.2021.118991
Caudullo, G., Vizzarri, M., & de Rigo, D. (2017). European Atlas of Forest Tree Species. European Commission, Joint Research Centre (JRC). https://doi.org/10.2760/36529
Carvalho, S. (2020). The effect of large tree removal on oak forest regeneration in Mediterranean landscapes. Ecological Indicators, 108, 105734. https://doi.org/10.1016/j.ecolind.2019.105734
Clotet, X., Alcañiz, M., & García, M. (2016). Invasive species in Mediterranean forests: The case of eucalyptus and pine plantations. Journal of Environmental Management, 179, 104- 112. https://doi.org/10.1016/j.jenvman.2016.04.014
Costa, J., Moreira, F., & Fernandes, P. (2021). Eucalyptus and pine plantations in cork oak habitats: Implications for forest biodiversity and management. Forestry, 94(3), 374-384. https://doi.org/10.1093/forestry/cpaa048
Davi, H., Gauthier, P., & François, P. (2020). Susceptibility of Mediterranean pine species to pathogens and climate stress. Journal of Vegetation Science, 31(2), 193-204. https://doi.org/10.1111/jvs.12810
Fernández-Manjarrés, J., Paredes, M., & Bonet, J. (2018). Aridity and nitrogen deposition effects on invasive Mediterranean woody species. Plant Ecology, 219(2), 145-159. https://doi.org/10.1007/s11258-017-0795-6
Fernández-Manjarrés, J., Bravo-Oviedo, A., & Martínez, M. (2022). Adaptive forest management under climate change: A case study of Mediterranean cork oak forests. Forest Ecology and Management, 510, 120076. https://doi.org/10.1016/j.foreco.2022.120076
García-Orenes, F., Bellot, J., & Martínez, A. (2022). Post-fire thinning effects on Mediterranean forest regeneration. Journal of Forestry Research, 33(2), 153-166. https://doi.org/10.1007/s11676-021-01370-4.
Gao, W., Wu, H., & Tian, S. (2019). The role of thinning in improving forest resilience to drought in Mediterranean ecosystems. Agricultural and Forest Meteorology, 271, 210-220. https://doi.org/10.1016/j.agrformet.2019.03.015
Gea-Izquierdo, G., Martínez-García, M., & Pinto, J. (2014). The impact of Phytophthora cinnamomi on different pine species in Mediterranean regions. Phytopathology, 104(3), 262- 273. https://doi.org/10.1094/PHYTO-07-13-0217-R
González, A., Muñoz, J., & Pino, J. (2020). The effects of drought and Phytophthora cinnamomi on oak forests in the Iberian Peninsula. Forest Pathology, 50(4), e12659. https://doi.org/10.1111/efp.12659
González-Moreno, P., Fernández-Rivas, M., & García, C. (2022). The role of tree species diversity in soil trophic networks and forest resilience. Ecological Applications, 32(5), e02504. https://doi.org/10.1002/eap.2504
Granados, J., Moreno, J., & Díaz-Fernández, P. (2021). Reforestation and afforestation for soil conservation and carbon sequestration in Mediterranean environments. Land Degradation & Development, 32(3), 1229-1243. https://doi.org/10.1002/ldr.3877
Hasnaoui, H., Messaoudi, F., & Mokhtar, M. (2021). Local population pressures and their impact on cork oak forests in Northwest Tunisia. Mediterranean Forestry Journal, 10(4), 62-75. https://doi.org/10.1080/09584240.2021.1904934
Helluy, S., Canellas, I., & Frey, A. (2023). Thinning effects on forest resilience in water-scarce environments. Forest Ecology and Management, 532, 120261. https://doi.org/10.1016/j.foreco.2022.120261
Ibañez, J., Martínez, A., & Ruiz-Peinado, R. (2022). Grazing management and wildfire risk reduction in Mediterranean forests. Journal of Environmental Management, 301, 113759. https://doi.org/10.1016/j.jenvman.2021.113759
IPCC. (2021). Climate Change 2021: The Physical Science Basis. Cambridge University Press. https://doi.org/10.1017/9781009157896
Jactel, H., & Brockerhoff, E. (2023). Tree diversity and forest resilience to disturbances: A review. Forest Ecology and Management, 522, 120356. https://doi.org/10.1016/j.foreco.2022.120356
Jactel, H., Tognini, S., & Dapporto, L. (2022). The influence of forest composition on pest dynamics and forest resilience. Journal of Applied Ecology, 59(8), 2267-2280. https://doi.org/10.1111/1365-2664.14150
Jaouadi, M., Rouini, N., & Haddad, H. (2021). Effects of thinning on pine tree growth and soil characteristics in Mediterranean environments. Plant and Soil, 460(1), 127-140. https://doi.org/10.1007/s11104-021-04800-x
Lindner, M., Zudin, S., & Cudlin, P. (2023). Forest responses to climate extremes: A global perspective. Global Change Biology, 29(1), 14-28. https://doi.org/10.1111/gcb.16305
Luque, J., Sánchez, M., & Martínez, M. (2002). Phytophthora cinnamomi and its effects on Quercus suber in Mediterranean ecosystems. European Journal of Forest Research, 121(1), 43-55. https://doi.org/10.1007/s10342-002-0008-2.
Marcais, B., Rizzo, D., & Lecomte, C. (2004). The effect of Phytophthora cinnamomi on soil biological activity and nutrient availability in Mediterranean oak forests. Soil Biology and Biochemistry, 36(7), 1151-1160. https://doi.org/10.1016/j.soilbio.2004.02.004
Martínez, M., & Fernández, S. (2022). Climate change impacts on forest ecosystems and forest management in the Mediterranean region. Forest Policy and Economics, 139, 102788. https://doi.org/10.1016/j.forpol.2022.102788
Mechergui, M., Ben Jabeur, M., & Boukerroui, S. (2022). Using Pinus halepensis as a rootstock for improved stone pine production in arid regions. Arid Land Research and Management, 36(1), 57-69. https://doi.org/10.1080/15324982.2021.1954371
Moralejo, E., Moricca, S., & Pérez, M. (2009). Susceptibility of Mediterranean oak species to Phytophthora cinnamomi. Forest Pathology, 39(5), 239-250. https://doi.org/10.1111/j.1439- 0329.2009.00590.x
Moricca, S., Garcia, E., & Rivas, J. (2016). Pathogen dynamics and host responses in Mediterranean oak forests under climate stress. Plant Pathology, 65(1), 86-100. https://doi.org/10.1111/ppa.12441
Moriondo, M., Giannakopoulos, C., & Bindi, M. (2022). Projections of forest fire risk under future climate scenarios in the Mediterranean Basin. Climate Change, 172(4), 439-456. https://doi.org/10.1007/s10584-022-03644-7
Muñoz-Rengifo, J., Díaz-Barradas, M., & González, A. (2021). Clearing shrublands to reintroduce Quercus suber in degraded Mediterranean ecosystems. Journal of Restoration Ecology, 29(2), 231-242. https://doi.org/10.1111/rec.13353
Olivar, J., Sabaté, S., & García, C. (2019). Effects of thinning on forest water use and growth under drought conditions. Hydrology and Earth System Sciences, 23(6), 2621-2634. https://doi.org/10.5194/hess-23-2621-2019
Pereira, P. (2007). Interaction between summer storms and Phytophthora cinnamomi in Mediterranean forest ecosystems. Plant Disease, 91(12), 1565-1571. https://doi.org/10.1094/PDIS-91-12-1565
Peñuelas, J., Canadell, J., & Lavorel, S. (2022). Grazing management and forest patch maintenance to stabilize Mediterranean ecosystems. Ecological Applications, 32(3), e02291. https://doi.org/10.1002/eap.2291
Pinto-Correia, T., & Godinho, S. (2020). The expansion of eucalyptus and pine plantations into cork oak habitats: Implications for biodiversity. Biodiversity and Conservation, 29(8), 2535- 2548. https://doi.org/10.1007/s10531-020-02072-3
Quinto, V., Morales, M., & Díaz, M. (2021). The role of seed-producing trees in the natural regeneration of cork oak forests. Forest Ecology and Management, 480, 118788. https://doi.org/10.1016/j.foreco.2020.118788
Rodríguez-Molina, M., Sánchez, J., & Moreno, G. (2002). Cork oak susceptibility to Phytophthora cinnamomi in Mediterranean ecosystems. Forest Pathology, 32(4), 252-264. https://doi.org/10.1046/j.1439-0329.2002.00282.x
Rosati, M., Vázquez, M., & Hernández, M. (2022). Sustainable cork harvesting and its effects on forest health and regeneration. Forest Management Review, 45(1), 22-33. https://doi.org/10.1016/j.fmr.2022.01.003
Ruiz-Peinado, R., García, R., & Martínez, S. (2022). The influence of shrub and grass reduction on tree planting success in Mediterranean forests. Plant Ecology, 223(5), 763-778. https://doi.org/10.1007/s11258-021-01196-6
Sánchez, M., Luque, J., & García, J. (2002). Cork oak decline in Mediterranean regions: Effects of Phytophthora cinnamomi and land management. Forest Pathology, 32(2), 119-136. https://doi.org/10.1046/j.1439-0329.2002.00209.x
Sánchez, R., González, A., & López, S. (2023). Extreme precipitation events and their impact on Mediterranean forest ecosystems. Hydrology and Earth System Sciences, 27(4), 1851-1865. https://doi.org/10.5194/hess-27-1851-2023
Sena, I., Oliveira, H., & Costa, A. (2018). Phytophthora cinnamomi and its impact on global forest composition changes. Forestry, 91(3), 263-278. https://doi.org/10.1093/forestry/cpx037
Tomaz, R., Ribeiro, S., & Monteiro, J. (2022). Reforestation and soil conservation practices in Mediterranean degraded areas. Land Degradation & Development, 33(2), 198-211. https://doi.org/10.1002/ldr.4186
Touhami, A., Zairi, M., & Bouaziz, A. (2024). Mitigating drought stress in cork oak forests through association with leguminous species. Journal of Arid Environments, 118, 56-67. https://doi.org/10.1016/j.jaridenv.2023.105745
Vilà-Cabrera, A., Moya, M., & Fernández, J. (2022). Enhancing water and carbon use efficiency in Mediterranean forests through thinning. Journal of Forest Research, 27(4), 417-429. https://doi.org/10.1007/s11676-022-01378-1
Wahbi, A., Bouanani, N., & Bouche, M. (2021). The effects of thinning on forest resilience to water scarcity in Mediterranean regions. Forest Ecology and Management, 484, 118887. https://doi.org/10.1016/j.foreco.2021.118887
Yachi, S., & Loreau, M. (2000). Biodiversity and ecosystem productivity in a fluctuating environment: The insurance hypothesis. Proceedings of the National Academy of Sciences, 97(11), 6579-6584. https://doi.org/10.1073/pnas.110534297.